Hetero version 2.0

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About this Collection

Hetero version 2.0

A program to simulate the evolution of DNA under the mixture models of heterogeneity across lineage and heterogeneity across sites. Hetero was written to facilitate the development and testing of phylogenetic methods. In particular, Hetero 2 allows the user to generate nucleotide sequences on a phylogenetic tree under complex conditions. The com... more

Bioinformatics Bioinformatics Software

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Hetero 2.0 Software Bioinformatics nucleotide sequences phylogenetic tree


Hetero code repository on GitHub


Vivek Jayaswal, Thomas K.F. Wong, John Robinson, Leon Poladian, Lars S. Jermiin; Mixture Models of Nucleotide Sequence Evolution that Account for Heterogeneity in the Substitution Process Across Sites and Across Lineages. Syst Biol 2014; 63 (5): 726-742. doi: 10.1093/sysbio/syu036



This software was developed by the bioinformatics and phylogenomics team in Ecosystem Sciences, The Commonwealth Scientific and Industrial Research Organisation (CSIRO), Canberra, Australia.

CSIRO Open Source Software Licence (Based on MIT/BSD Open Source Licence)

CSIRO (Australia)

Jayaswal, Vivek; Wong, Thomas KF; Robinson, John; Poladian, Leon; Jermiin, Lars (2014): Hetero version 2.0. v1. CSIRO. Software Collection.

All Rights (including copyright) CSIRO 2014.

The metadata and files (if any) are available to the public.

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More about this Collection

Tested on Linux and MacOS




About this Project

CLSD Strategic: Solving Biodiversity Cha

We generate knowledge on the mechanistic links and synergistic interactions between landscape change, species invasions and native species resilience in terrestrial ecosystems. Working on both natural and agricultural systems, we undertake research and develop theory to underpin more effective policy and management actions for conservation, invasio... more

Bruce Webber

Bioinformatics Software

Bioinformatics software previously made available from the CSIRO ICT and Services website.


Vivek Jayaswal

Thomas KF Wong

John Robinson

Leon Poladian

Lars Jermiin

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